First description of the early Devonian ammonoid Mimosphinctes from Gondwana and stratigraphical implications

Swiss Journal of Palaeontology

Table 4 Ammonoid stratigraphy of the Emsian Stage of the eastern Anti-Atlas (Morocco)

Stage	Lithostratigraphic units (beds)	Index ammonoids	Additional ammonoids
Late Emsian	Upper Anarcestes Lst.	Anarcestes lateseptatus	Amoenophyllites doeringi, Anarcestes spp., Chlupacites praeceps, Mimagoniatites bohemicus
	Lower Anarcestes Lst. (Sellanarcestes Marls)	Sellanarcestes wenkenbachi	Achguigites tafilaltensis, Anarcestes spp., Chlupacites praeceps, Paranarcestes pictus, Sellanarcestes neglectus
	Middle and upper Daleje Shale	Latanarcestes noeggerathi auct.	Latanarcestes latisellatus, Rherisites tuba, Praewerneroceras hollardi
	Lower Daleje Shale	Gyroceratites gracilis	Rherisites tuba ^a
	Base Daleje Shale	Gyroceratites laevis	–
Early/late Emsian	Marls above Mimagoniatites Lst.	Mimosphinctes karltschanzi	–
Early Emsian	Mimagoniatites Lst.	Mimagoniatites fecundus	–
	Upper Erbenoceras Lst.	Klugites gesinae	Erbenoceras solitarium, Weyeroceras angustum
	Lower Erbenoceras Lst.	Anetoceras obliquecostatum	Anetoceras obliquecostatum, Erbenoceras advolvens, Irdanites korni, Metabactrites formosus, Teicherticeras cf. senior
	Lower Devonobactrites Shale (early Faunule 2)	Erbenoceras advolvens	Anetoceras sp., Irdanites kaufmanni, Ivoites ernsti, Metabactrites formosus, Gyroceratites cf. heinricherbeni, Chebbites spp.
	Deiroceras Lst.	Chebbites reisdorfi	–

This table was assembled using own field data as well as published data from Becker and House (1994), Klug (2001, 2002), Klug et al. (2008), De Baets et al. (2010), Aboussalam et al. (2015) and this paper. It is unclear whether the Gyroceratites laevis, G. gracilis and Latanarcestes noeggerathi (auct.) zones are entirely justified and to which degree the occurrences overlap. In that respect, this table is an attempt to summarize the state of research; at the same time, the lack of biostratigraphic data have to be stressed as well as the need for further research
^aAboussalam et al. (2015, p. 926) noted that Rherisites tuba enters in the Daleje Shale Equivalent before the first anarcestids (W of Hamar Laghdad), which justifies a new Rh. tuba Zone. They also emphasised that the true Lat. noeggerathi is a younger species (holotype from the Wissenbach Slate) than the oldest anarcestids commonly given that name